Combined quantum mechanical/molecular mechanical (QM/MM) methods

 

·     To treat a chemical reaction, a quantum mechanical method is required to describe the breaking and forming of bonds - such methods include ab initio and semiempriical molecular orbital methods. 

 

·     Molecular mechanics methods are not suitable, as outlined above. 

 

·     However, ab initio methods cannot be applied to systems with more than a few tens of atoms, because the computational demands are too great. 

 

·     Semiempirical methods can be applied to bigger systems (100s of atoms), but even they are too demanding for calculations on very large systems. 

 

·     For calculations on reactions in solution, where thousands of solvent molecules may need to be included, or for reactions in solids, at surfaces or in biological macromolecules (e.g. enzymes), a different approach must be used. 

 

·     Quantum mechanical/molecular mechanical (QM/MM) methods allow reactions in large systems to be studied. 

 

·     A small region including the reacting atoms is treated with a QM method (e.g. ab initio molecular orbital theory), while most of the system is represented more simply by molecular mechanics.  

 

·     The QM (e.g. reacting) region ‘feels’ the influence of the molecular mechanics environment - for example, the atomic charges of the MM atoms affect the QM atoms. 

 

·     By combining the accuracy of a QM method (for a small reacting system) with the speed of a MM method (for its surroundings), reactions in large molecules and in solution can be studied. 

Schematic representation of the setup of a QM/MM calculation.

A small region containing the reacting atoms is treated by a quantum mechanical method (e.g. molecular orbital theory), and its surroundings are represented more simply by molecular mechanics.

 

The total energy of the system can be written as:

 

E= E(QM) + E(QM/MM) + E(MM) + E(boundary)

 

·     The energy of the QM system, E(QM), and the energy of the MM system, E(MM) are calculated exactly as they would be in a standard calculation at those levels.

 

·     E(QM/MM) is the interaction energy between the QM and MM regions (e.g. due to the interaction of the MM atomic charges with the QM system)

 

·     E(boundary) the energy due to any boundary restraints applied to the outer edge of the MM region to maintain its structure.

 

 

Modelling an enzyme-catalysed reaction with a QM/MM method. 

The picture shows the active site of para-hydroxybenzoate hydroxylase, which hydroxylates (adds an OH group) to its substrate, para-hydroxybenzoate.  The atoms treated by QM are shown as (thick) bonds. Other groups in the enzyme are treated MM in the calculation (some of the important surrounding residues are shown in thin lines). The transition state for hydroxylation is shown (Od is being transferred from the flavin cofactor on the right, to form a new C-O bond with the substrate (shown on the left).  

 

Introduction to molecular mechanics